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At each amino acid position the mutation rate from one charge state to the other is \\(\\nu\\). We derive the expected variance in charge, for a population of N individuals in generation t, which is \\(\\sigma^ 2(t)=[2MN\\nu /(1+8N\\nu)]\\{1-\\exp [- (4\\nu +1/(2N))t]\\}\\) for an initially monomorphic population.    This variance, which is unaffected by intragenic recombination and gene conversion, does not increase linearly with N as does the steady-state variance in charge for the older electrophoretic models. We also derive the probability, \\(P_ j(t)\\), that a random gene in a population exceeds another by j charge units.    We conclude: (1) that, except for species which have maintained effective population numbers in the millions, the infinite mutable sites model adequately describes neutral genetic variation; (2) that very high levels of 'neutral' genetic variation can be attained only by genes with a fairly large number of 'neutral' amino acid sites in species which have maintained an effective size in the millions for millions of generations; (3) that, because all species surveyed have a mean heterozygosity of less than 25\\%, it is likely that most genes have only a few 'neutral' 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