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By applying the continuation theorem in \\textit{R. E. Gaines} and \\textit{J. L. Mawhin} [Coincidence degree, and nonlinear differential equations. Lecture Notes in Mathematics. 568. Berlin etc.: Springer-Verlag (1977; Zbl 0399.47031)], sufficient conditions are derived for the existence of positive periodic solutions to (1). The results of this paper give an affirmative answer to the open problem 9.1 due to \\textit{Y. Kuang} [Delay differential equations: with applications in population dynamics. Mathematics in Science and Engineering. 191. Boston, MA: Academic Press Inc. (1993; Zbl 0777.34002)]. Some existing results due to \\textit{K. Gopalsamy}, \\textit{X. He} and \\textit{L. Wen} [Glasg. Math. J. 33, 281--286 (1991; Zbl 0737.34050)], \\textit{H. Fang} and \\textit{J. Li} [J. Math. Anal. Appl. 259, 8--17 (2001; Zbl 0995.34073)] and \\textit{Li Yong Kun} [J. Math. Anal. Appl. 214, 11--21 (1997; Zbl 0894.34075)] are extended and 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