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The paper is concerned with the investigation of the harmonicity of some special endomorphisms and semi-Riemannian structures in Walker geometry in four dimensions. The authors first establish a one-to-one correspondence between the class of harmonic Walker metrics, with respect to a given Walker metric, and a certain transformation group. Then, they characterize the harmonicity of the Sasaki, horizontal and vertical lifts of a Walker metric in terms of the harmonicity of the Walker metric on the base manifold. They also deal with the almost hyper-para-Hermitian structure defined by two almost para-complex structures and one almost complex structure, satisfying certain conditions. In this context, they prove that the two almost para-complex structures are harmonic if and only if they are almost para-K\u00e4hler, and similarly for the almost complex structure. 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