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The authors exploit a map of classifying spaces \\(BS^1\\to BS^3\\) so as to relate the systolic ratios of the quaternionic projective space and the complex projective space, and to relate the quaternionic projective space and a hypothetical Joyce manifold (with Spin(7)-holonomy) with \\(b_4=1\\). They study optimal curvature-free inequalities of the type discovered by C.~ Loewner and M.~Gromov, using a generalization of the Wirtinger inequality for the comass. Using a model for the classifying space \\(BS^3\\) built inductively out of \\(BS^1\\), the authors prove that the symmetric metrics of certain two-point homogeneous manifolds turn out not to be the systolically optimal metrics on those manifolds. By calculating the Wirtinger constants for \\(\\mathbb R^8\\) the authors show that the common value of the optimal systolic ratio for \\(\\mathbb H\\mathbb P^2\\) and \\(\\mathbb C\\mathbb P^4\\) satisfies \\(6\\leq \\text{SR}_4(\\mathbb H\\mathbb P^2) = \\text{SR}_4(\\mathbb C\\mathbb P^4)\\leq 14\\). The constant \\(14\\) is twice the dimension of the Cartan subalgebra of the exceptional Lie algebra 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