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    1986
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    diploid one-locus two-allelic diffusion model for group selection
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    infinite number of demes
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    mutation
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    random genetic drift
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    migration
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    altruistic trait
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    Kolmogorov forward diffusion equation
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    population genetics
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    interdeme competition
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    Wright's island model
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    semi-dominance selection scheme
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    approximate steady-state distribution
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