Permanence of periodic predator-prey system with two predators and stage structure for prey (Q1049466)

The authors investigate a four-dimensional nonautonomous predator-prey system with two predators, and a prey divided into an immature and a mature population. The righthand side is time-periodic and contains functional responses of type Holling IV and Beddington-DeAngelis. The aim of the paper is to establish a set of sufficient and necessary conditions that guarantee the permanence of the system. (The proof of necessity seems to be incomplete.) Two concluding examples illustrate the theoretical results.