External stability and ESS: Criteria for initial increase of new mutant allele (Q750352)

\textit{I. Eshel} and \textit{M. W. Feldman} [Am. Natur. 124, 631-640 (1984)] developed an average fitness condition for the initial increase of a new mutant allele from a stable equilibrium for the case of two multiple- allelic loci, such that the mutant allele succeeds only when its average marginal viability exceeds that of the resident population. In other words, the original stable equilibrium is externally stable only when its average fitness exceeds that associated with the new mutant allele. We generalize this fitness condition and show that it applies to the more general framework of multilocus multiallele viability selection models.