Existence of positive periodic solution of a periodic cooperative model with delays and impulses (Q5971236)

A periodic cooperative species model with delays and impulses, \[ \begin{aligned} \frac{dx_1(t)}{dt} = r_1(t)x_1(t)\Big [ 1-\frac {x_1(t-\tau_{11}(t))}{a_1(t)+b_1(t)x_2(t-\tau_{12}(t))} \\ -c_1(t) x_1(t-\tau_{13}(t))\Big], \qquad t>0, t\neq t_k, \\ \Delta x_1(t_k) = -\gamma_{1k}x_1(t_k), \qquad k=1,2, \dots, \\ \frac {dx_2(t)}{dt} = r_2(t)x_2(t)\Big [1-\frac{x_2(t-\tau_{21}(t))}{a_2(t)+b_2(t)x_1(t-\tau_{22}(t))} \\ -c_2(t)x_2(t-\tau_{23}(t))\Big], \qquad t>0, t\neq t_k, \\ \Delta x_2(t_k) = -\gamma_{2k}x_2(t_k), \qquad k=1,2, \dots, \end{aligned} \] is proposed and studied. The model generalizes the one proposed by [\textit{R.\ M.\ May}, Stability and Complexity in Model Ecosystems Princeton Landmarks in Biology. (Princeton), NJ: Princeton University Press (2001; Zbl 1044.92047)] by incorporating periodically changing environment and impulsive birth. The existence of at least one positive periodic solution to the model is established by using the continuation theorem of coincidence degree theory.